Dr. Dale Tshudy, Professor, Geosciences, is a geologist/paleontologist who joined the Edinboro University faculty in 1993. His research interests are in the fossil record of crustaceans, especially lobsters, but he has taught a broad array of geology and oceanography courses at Edinboro University and at the Wallops Island (Virginia) Marine Science Consortium's field station.
Department of Geosciences
Edinboro University of Pennsylvania
Edinboro, Pennsylvania 16444
(814) 732-2453 (office - Cooper Hall 235)
(814) 732-2529 (Geosciences main office)
Office hours for Spring, 2013: T 2-3; W 11-12, 1-2; R 2-3; F 1-2; also by appointment.
Ph.D., Geology, Kent State University, 1993. Dissertation title: "Taxonomy and Evolution of the Clawed Lobsters, Families Nephropidae and Chilenophoberidae" (472 p.).
M.S., Geology, Kent State University, 1987. Thesis title: "Macruran Decapods, and their Epibionts, from the Lopez de Bertodano Formation (Late Cretaceous), Seymour Island, Antarctica".
B.S., Geology, minor in Geography, University of Delaware, 1985.
Teaching Interests (Courses offered at Edinboro University main campus listed below with GS prefix; courses offered at Wallops Island Marine Science Consortium's field station listed below with "MS" prefix).
Dynamic Earth (physical geology) (GS 101)
Introduction to Earth Environments (introductory environmental geology) (GS 104)
Principles of Earth History (GS 112)
Field Methods in Oceanography (MS 211)
Introduction to Oceanography (GS 310 / MS 310)
Marine Geology (MS 362)
Environmental Geology (upper level) (GS 375)
Independent Study (invertebrate paleontology, vertebrate paleo. (GS 490)
Coral Reef Ecology (1 week at Wallops, 2 in Roatan, Honduras) (MS 491)
Regional Geology (GS 522)
Invertebrate Paleontology (GS 523)
Field Studies in the Geosciences (GS 525)
Seminar in the Geosciences (a field course in limnology) (GS 595)
Crustacean (especially clawed lobster) evolution and taxonomy as interpreted by both morphologic and molecular methods; also taphonomy (preservation), biogeography, paleoecology and extinction.
Current (updated 3-13) research interests include: 1) describing new fossil lobster species from various locales/strata; 2) morphologic and molecular (DNA-based) analyses of clawed lobster phylogeny (evolutionary relationships); 3) aspects of clawed lobster biogeography.
Publications in Refereed Journals
Tshudy, D. and G. A. Bishop, in preparation. New fossil clawed lobsters (Family Nephropidae) from the Carlile Shale (Cretaceous: Turonian), South Dakota.
Tshudy, D. and U. Sorhannus, in preparation. Phylogeny of the clawed lobsters, families Chilenophoberidae, Nephropidae and Thaumastochelidae: analysis of an expanded morphology matrix.Tshudy, D. and J. Saward. 2012. Dinochelus steeplensis, a new species of clawed lobster (Nephropidae) from the London Clay (Eocene) of England. Journal of Crustacean Biology 3291):67-79.
De Grave, S. et al. 2009. A Classification of Living and Fossil Genera of Decapod Crustaceans. The Raffles Bulletin of Zoology, Supplement 21, 93 p.
Tshudy, D., T-Y Chan, and U. Sorhannus. 2007. Morphology-based cladistic analysis of Metanephrops, the most diverse extant genus of clawed lobster (Family Nephropidae). Journal of Crustacean Biology 27(3):463-476.
Tshudy, D., S. Donaldson, C. Collom, R. M. Feldmann and C.E. Schweitzer. 2005. Hoploparia albertaensis, a new species of clawed lobster (Nephropidae) from the late Coniacian, shallow-marine Bad Heart Formation of northwestern Alberta, Canada. Journal of Paleontology, 79(5):961-968.
Tshudy, D. and U. Sorhannus. 2003. Hoploparia, the best-known fossil clawed lobster (Family Nephropidae) is a wastebasket genus. Journal of Crustacean Biology, 23(3):712-722.
Tshudy, D. 2003. Clawed lobster (Nephropidae) diversity through time. Journal of Crustacean Biology, 23(1):178-186.
Tshudy, D. and U. Sorhannus. 2000. Jagtia kunradensis, a new genus and species of clawed lobster (Decapoda: Nephropidae) from the Upper Cretaceous (Upper Maastrichtian) Maastricht Formation, The Netherlands. Journal of Paleontology, 74(2):224-229.
Tshudy, D. and U. Sorhannus. 2000. Pectinate claws in decapod crustaceans: convergence in four lineages. Journal of Paleontology, 74(3):474-486.
Tshudy, D., Baumiller, T. K. and U. Sorhannus. 1998. Morphologic change in the clawed lobster Hoploparia from the Cretaceous of Antarctica. Paleobiology, 24(1):64-73.
Tshudy, D. and G. Parsons. 1998. Intraspecific variation in the Maine lobster, Homarus americanus. Proceedings of the Biological Society of Washington, 111(1): 102-109.
Tshudy, D. and L. E. Babcock. 1997. Morphology-based phylogenetic analysis of the clawed lobsters, Family Nephropidae and Family Chilenophoberidae. Journal of Crustacean Biology, 17(2):253-263.
Gryta, J. J., D. Tshudy and J. F. Lichtinger. 1995. Bedrock geology of the Edinboro North 7 1/2' Quadrangle, Erie County, Pennsylvania. Open File Report 95 of the Pennsylvania Geological Survey.
Feldmann, R. M., D. M. Tshudy, and M. A. Thomson. 1993. Cretaceous-Paleocene decapod crustaceans from the James Ross Basin, Antarctic Peninsula. Paleontological Society Memoir Series, 28.
Jagt, J. R. M. Feldmann, and D. M. Tshudy. 1990. Oncopareia bredai from the lower upper Maastrichtian of Maastricht. Mededelingen van der Rijks Geologische Diesnt, 44(1):23-35.
Gibbs, R. J., D. M. Tshudy, L. Konwar, and J. Martin. 1989. Coagulation and transport of sediments in the Gironde Estuary, France. Sedimentology, 36(5):621-626.
Tshudy, D. M., R. M. Feldmann, and P. D. Ward. 1989. Cephalopods: biasing agents in the preservation of lobsters. Journal of Paleontology, 63(5):621-626.
Feldmann, R. M. and D. Tshudy. 1987. Ultrastructure in cuticle from Hoploparia stokesi (Decapoda: Nephropidae) from the Lopez de Bertodano Formation (Late Cretaceous-Paleocene) of Seymour Island, Antarctica. Journal of Paleontology, 61(6):1194-1203.
Tshudy, D. in press. Chapter 1, “Systematics”, in “Ecology and Biology of the Norway Lobster, Nephrops norvegicus” (Magnus Johnson and Mark Johnson, eds). Elsevier Press, Oxford.
Wahle, R. A., D. Tshudy, J.S. Cobb, J. Factor and M. Jaini. 2012. Traite de Zoologie, Chapter 66 (pp.3-108): Infraorder Astacidea Latreille, 1802 P.P..: The Marine Clawed Lobsters. Leiden, Brill Publishing.Tshudy, D., R. Robles, T.-Y. Chan, K. C. Ho, K. H. Chu, S. T. Ahyong and D. Felder, 2009. Phylogeny of marine clawed lobsters families Nephropidae Dana, 1852, and Thaumastochelidae Bate, 1888, based on mitochondrial genes. In: Martin, J. W., K. A. Crandall and D. L. Felder, Decapod Crustacean Phylogenetics. Koenemann, S., Crustacean Issues 18. Boca Raton, London, New York, CRC Press, Taylor and Francis Group, Pp. 357-368.
Patek, S. N., R. M. Feldmann, M. Porter, and D. Tshudy. 2006. Phylogeny and Evolution. In: B. F. Phillips, Lobsters: Biology, Management, Aquaculture and Fisheries. Blackwell Publishing.Feldmann, R. M., and D. M. Tshudy. 1989. Evolutionary patterns in macrurous decapod crustaceans from Cretaceous to early Cenozoic rocks of James Ross Basin, Antarctica. In J. A. Crame (ed.), Origins and Evolution of the Antarctic Biota, Geological Society of London Special Publication 47:183-195.
Tshudy, D. M. and R. M. Feldmann. 1988. Macruran decapods, and their epibionts, from the Lopez de Bertodano Formation (Upper Cretaceous), Seymour Island, Antarctica. In R. M. Feldmann and M. Woodburne (eds.), Geology and Paleontology of Seymour Island, Antarctica, Geological Society of America Memoir 169:291-301.
Published Abstracts for Papers Presented by Tshudy (excludes abstracts presented by co-authors)
Tshudy, D. and U. Sorhannus, 2011. A cladistic analysis of clawed lobsters based on an expanded morphological data set. Abstracts with programs. Geological Society of America. Northeastern/North-Central Section Meeting. Pittsburgh, PA.
Tshudy, D. , S. Ahyong, T-Y Chan, U. Sorhannus. 2008. An expanded, morphology based cladistic analysis of the clawed lobsters. Society for Integrative and Comparative Biology 2008 Annual Meeting, San Antonio, Texas.
Tshudy, D., T.-Y. Chan, U. Sorhannus. 2006. Morphology-based cladistic analysis of Metanephrops, the most diverse extant genus of clawed lobster (Nephropidae). Abstracts with Programs, Geological Society of America, North-Central Section Meeting, Akron, Ohio.
Tshudy, D., K. H. Chu, R. Robles, K. C. Ho, T.-Y. Chan, S. T. Ahyong, and D. Felder. 2005. Phylogeny of marine clawed lobsters based on mitochondrial RDNA. Sixth International Crustacean Congress (Glasgow, Scotland) Abstracts O6.
Tshudy, D. and L. Babcock. 1995. A morphology-based cladistic analysis of the clawed lobsters (Family Nephropidae). Joint Meeting of the American Society of Zoologists and the American Crustacean Society, American Zoologist, 35(5):14A.
Tshudy, D. M. and R. M. Feldmann. 1991. Rates and modes of evolution in Late Cretaceous-Early Tertiary nephropid lobsters (Crustacea:Decapoda) from James Ross Basin, Antarctica. Geological Society of America, Abstracts with Programs, 23(5):405.
Feldmann, R. M. and D. M. Tshudy. 1990. Elongate claws in decapod crustacean taxa; "preadaptation" to scavenging in aphotic habitats. Geological Society of America, Abstracts with Programs, 22(5):10.
Tshudy, D. M., R. M. Feldmann, and P. D. Ward. 1988. Scavenging of lobster remains by Nautilus, and its taphonomic implications. Geological Society of America, Abstracts with Programs, 20(5):10.
Tshudy, D. M. and R. M. Feldmann. 1987. Taphonomy of decapods from the Lopez de Bertodano Formation (Cretaceous), Antarctic Peninsula. Geological Society of America, Abstracts with Programs, 19(4):249-250.
Tshudy, D. M. and R. M. Feldmann. 1986. Macrurous decapod crustaceans from the Lopez de Bertodano Formation (Late Cretaceous), Seymour Island, Antarctica. Geological Society of America, Abstracts with Programs, 18(4):328.
Gibbs, R. J., D. Tshudy, L. Konwar, and J. Martin. 1985. Suspended sediment transport of the Gironde Estuary, France. Geological Society of America, Abstracts with Programs, 22(5):10.
Citations in Popular Press
Corson, T., 2004, The Secret Life of Lobsters, p.165 (re: lobster diversity through time)
Morris, S. C., 2003, Life's Solution: Inevitable Humans in a Lonely Universe, p.130, 363 (re: convergent evolution)
Selected newspaper articles
“Lobster lowdown; Edinboro prof studies likely missing link”. Erie Times-News, January 24, 2011. P. 3B. Article by Sean McCracken. Story ran in AP. Original release, written by Ursula Wheeler, ran as “’Missing Link’ Lobster” in Meadville Tribune on Feb. 1, 2011, p. A8.
“A reputation evolved; How we came to be”. Erie Times-News, February 12, 2009. Article by Cody Switzer.
“Edinboro scientists join forces; Grant lets faculty, students compare methods as well as critters, professor says”. Erie Times-News, Summer (date?), 1996. Article by Tim Hahn.
“Of fossils and philosophies; Lobsterman aids academic in evolutionary research”. The Bar Harbor Times, June 22, 1995, p. B1. Article by Nancy Richards.
“Edinboro geologists discover giant prehistoric fish in county”. Erie Times-News, September 11, 1994. Article by Brian Pitzer.
Tshudy, 2008, $2013 awarded by Edinboro University Faculty Senate Research Committee for purchase of photographic equipment for paleontology research.
Summer 2007, collaborating investigator in the National Science Foundation's "Assembling a Tree of Life" (decapod crustaceans) intitiative (supplemental grant to Keith Crandall, NSF Award Abstract #0531762); summer stipend to support morphologic analysis of decapod crustacean evolution.
Summer 2006, collaborating investigator in the National Science Foundation's "Assembling a Tree of Life" (decapod crustaceans) intitiative (supplemental grant to Darryl Felder, NSF Award Abstract #0531603); summer stipend to support morphologic and molecular (DNA) analyses of decapod crustacean evolution.
Tshudy, 2006, "Initiation of a Crustacean DNA Sequencing Program at Edinboro University"; $2500 awarded by Edinboro University Faculty Senate Research Committee for laboratory supplies and to support collection of materials for DNA sequencing at the Smithsonian Institution.
Tshudy, 2005, "Morphology-based analysis of clawed lobster evolution"; $1445 awarded by Edinboro University Faculty Senate Research Committee to support travel to the Smithsonian Institution.
NSF RUI:ROA [National Science Foundation, Research at Undergraduate Institutions: Research Opportunity Award (Amendment No. 001 to Grant No. 0315995 to S. Fredericq and D. Felder)]; $11,371 to support "Molecular Phylogeny of the Clawed Lobsters, Family Nephropidae" at Dr. Darryl Felder's Laboratory for Crustacean Research, University of Louisiana at Lafayette, in Fall, 2004.
Tshudy and Van Bell, 1997, "Molecular Phylogeny of the Clawed Lobsters"; awarded $3,665 by State System of Higher Education (SSHE) Faculty Development Council.
Tshudy, Sorhannus, Fox and Van Bell, 1997, "Studies in Molecular Phylogeny"; $2,000 awarded by Edinboro University (President's Faculty Development Funds).
Tshudy, 1997, "Molecular Phylogeny of the Clawed Lobsters"; $920 awarded by Edinboro University Faculty Senate Research Committee.
Sorhannus, Van Bell and Tshudy, 1996, "Molecular Phylogeny"; $20,579 awarded by National Science Foundation. Edinboro University matched this award. The grant has enabled us to purchase a variety of equipment/materials necessary for student and faculty research on DNA/RNA-based methods for interpreting evolutionary relationships.
Tshudy, 1995. "An Analysis of Intraspecific and Ecophenotypic variation in the Clawed Lobster, Homarus americanus"; $1,978 awarded by Edinboro University Faculty Senate Research Committee.
Gryta and Tshudy, 1994, "Bedrock Geology of the Edinboro North Quadrangle, Erie Co., Pa."; awarded $10,000 by the Pennsylvania Geological Survey.
Tshudy, 1993, "A Cladistic Analysis of the Clawed Lobsters"; $225 awarded by Edinboro University Senate Faculty Research Committee.
Tshudy, 1991, "Taxonomy and Evolution of the Clawed Lobsters, Family Nephropidae"; $600.00 awarded by the Lerner-Gray Fund for Marine Research.
Awards and Honors
2002 – Faculty Researcher of the Year Award, Edinboro University of Pennsylvania
1999 – Presidential Research Award, Edinboro University of Pennsylvania
1992 – Kent State University “David B. Smith Award” for outstanding scholarship and research potential
1987 – Sigma Xi, KSU Chapter, “Outstanding Graduate Research Award”